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Study
A study (project) groups together data submitted to the archive and controls its release date. A study accession is typically used when citing data submitted to ENA. Note that all associated data and other objects are made public when the study is released.
Sample
A sample contains information about the sequenced source material. Samples are associated with checklists, which define the fields used to annotate the samples. Samples are always associated with a taxon.
Experiment
An experiment contains information about a sequencing experiment including library and instrument details.
Run
A run is part of an experiment and refers to data files containing sequence reads.